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photograph
Mature tree growing in the mountains N of Los Angeles, CA [C.J. Earle, 8-Mar-1998].

photograph

Bark on a tree in the White Mountains, California. Width of view about 30 cm [C.J. Earle, 27-Mar-1987].

photograph

Foliage and seed-bearing cones on a tree in the White Mountains, California. Width of view about 15 cm [C.J. Earle, 27-Mar-1987].
Pinus monophylla Torrey & Frémont 1845

Common Names

Singleleaf piñon (3); one-leaved, Gray, Frémont, or Nevada nut pine (5).

Taxonomic notes

Subsection Cembroides (6). Syn: Caryopitys monophylla (Torrey & Frémont) Rydberg; Pinus californiarum D.K. Bailey; P.cembroides Zuccarini var. monophylla (Torrey & Frémont) Voss (1).

Description

Trees 5-9(14) m tall and up to 50 cm diameter, strongly tapering, much branched; crown usually rounded, dense. Bark red-brown, irregularly furrowed or cross-checked, scaly. Branches spreading and ascending, persistent to near trunk base; twigs stout, orange-brown, aging brown to gray, sometimes sparsely puberulent. Buds ellipsoid, light red-brown, 0.5-0.7 cm, resinous; scale margins fringed. Needles 1 (rarely 2) per fascicle, ascending, persisting 4-6(10) years, 2-6 cm x 1.3-2(2.5) mm, stout, curved, terete (though often 2-grooved), gray-green, all surfaces with stomatal lines, margins entire, apex subulate; sheath 0.5-1 cm, scales soon recurved, forming rosette, shed early. Staminate cones ellipsoid, ca. 10 mm, yellow. Ovulate cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, ovoid before opening, broadly depressed-ovoid to nearly globose when open, 4-6(8) cm, pale yellow-brown, nearly sessile; apophyses thickened, slightly raised; umbo subcentral, raised or depressed, nearly truncate, apiculate. As with other piñons, the seeds rest in a deep cone-scale declivity and upper cone scale tissue holds the seeds in place, so seeds do not readily fall out and are readily available to avian dispersers. Seeds cylindric-ellipsoid; body 15-20mm, gray-brown to brown, wingless, edible. 2 n =24 (7, 9, 12).

Range

USA: Idaho, Utah, Nevada, Arizona, California; Mexico: Baja California Norte. 1000-2300 m. Prefers dry, gravelly slopes in semiarid country, where it forms woodlands alone or with juniper ( Juniperus sp.) (7, 9). See also (13).

Big Tree

Diameter 133 cm, height 14 m, crown spread 12 m, located in Inyo County, California (11).

Oldest

652 years. This is based on the Pequop Summit tree-ring chronology collected by Charles Stockton, with one core from a presumably live tree, dating from 1330 to 1982 (10).

Dendrochronology

Has been used in stable-isotope studies, historical archeology (dating the construction of old mine buildings), air pollution assessment, and a wide variety of climate studies (8). NOAA lists 15 chronologies for this species (10).

Ethnobotany

The large, nutritious seeds were a staple food for native americans living within the range of this species.

Observations

Seen widely throughout its range. Piñon-Juniper woodlands are particularly well-developed in east-central Nevada, while the largest and most impressive individual trees I have seen have been in Joshua Tree National Park (CA), near the southwest range limits of the species.

Remarks

The only pine bearing a single needle per fascicle.

Hybridizes with P.edulis and P.quadrifolia (9).

Singleleaf piñon is the state tree of Nevada (9).

Citations

(1) Silba 1986 .

(2) Jaeger .

(3) Arno & Gyer 1973 .

(5) Peattie 1950 .

(6) Perry 1991 .

(7) Little 1980 .

(8) Bibliography of Dendrochronology .

(9) Robert Kral in the Flora of North America online .

(10) NOAA Tree Ring Search Page .

(11) American Forests 1996 .

(12) Ronald M. Lanner, e-mail communication, 20-Dec-1999.

(13) Robert S. Thompson, Katherine H. Anderson and Patrick J. Bartlein. 1999. Atlas of Relations Between Climatic Parameters and Distributions of Important Trees and Shrubs in North America. U.S. Geological Survey Professional Paper 1650 A&B. URL= http://greenwood.cr.usgs.gov/pub/ppapers/p1650-a/pages/conifers.html, accessed 22-Jan-2000.

See also:

Burns & Honkala 1990 .

Farjon & Styles 1997 .

FEIS database .

Lanner 1981 .

Lanner 1983 .

Muir 1894 .


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This page is from the Gymnosperm Database
URL: http://www.geocities.com/RainForest/Canopy/2285/pi/pin/monophylla.htm
Edited by Christopher J. Earle
E-mail: earlecj@earthlink.net
Last modified on 28-Jan-2000

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